Posts Tagged ‘in tandem’
May 31, 2019
An Aurora Damsel (Chromagrion conditum) was spotted along the shoreline of a small pond located in Prince William County, Virginia USA. Aurora Damsel is a new species for my life list odonates.
This individual is a male, as indicated by his coloration and terminal appendages. Speaking of coloration, notice the underside of the male’s thorax is yellow — a key field mark for Aurora.
A mating pair of Aurora Damsel was spotted at the same location. This pair is “in tandem“: the male is on the upper-right; the female on the lower-left.
After copulation, Aurora Damsel engages in a form of guarding behavior known as “contact guarding,” in which the male and female fly “in tandem” to egg-laying sites. Contact guarding is used by most species of damselflies and some species of dragonflies to prevent aggressive males from hijacking the female.
Female C. conditum is polymorphic, including two morphs: blue coloration similar to male; or with an entirely yellow thorax, as shown above.
It’s helpful to take photos of mating pairs of damselflies, especially “in tandem,” since males and females of the same species can look quite different. The preceding photo is slightly overexposed. As a result, the yellow coloration on both the male and female looks a little washed out.
Credits
Sincere thanks to Gary Myers for the tip that enabled Mike Powell and me to find this uncommon damselfly. See Aurora Damsels in action for Mike’s take on our first time seeing this species.
Copyright © 2019 Walter Sanford. All rights reserved.
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Tags:21 MAY 2019, adult female, adult male, Aurora Damsel, Chromagrion conditum, Family Coenagrionidae, in tandem, mating pair, Narrow-winged Damselflies, polymorphic, terminal appendages
Posted in Aperture, Canon 430EX II Speedlite, damselflies, digital photography, Lightroom, natural science, Panasonic DMC-FZ300, Photoshop, wildlife photography | 2 Comments »
August 5, 2018
A mating pair of Stream Bluet damselflies (Enallagma exsulans) was spotted during a stream-walk along South Fork Quantico Creek in Prince William Forest Park (PWFP), Prince William County, Virginia USA. This pair is in tandem.
After copulation, Stream Bluet engages in a form of guarding behavior known as “contact guarding,” in which the male and female fly “in tandem” to egg-laying sites. Contact guarding is used by some species of odonates to prevent aggressive males from hijacking the female.
Female E. exsulans is polymorphic, including two morphs: green or blue thorax; abdomen similar to male for both morphs. The female shown in the preceding photo is a green morph.
Look closely at the posterior end of the female’s abdomen. All female damselflies (and some species of dragonflies) use an ovipositor to insert fertilized eggs into vegetation (endophytic oviposition). Notice the white spheroid at the tip of her ovipositor. That’s either a single egg or egg cluster/mass, probably the former.
Now look at the male. He appears to be “recharging” for mating again: Sperm is transferred from the genital opening under abdominal segment nine (S9) to the secondary genitalia located under abdominal segment two (S2). Remember all dragonflies and damselflies have a 10-segmented abdomen.
Editor’s Notes
Thanks to my good friend Mike Boatwright, administrator of the Virginia Odonata Facebook group, for verifying my tentative identification of the species of damselfly. Also thanks to Sue Gregoire, Kestrel Haven Migration Observatory, for sharing her expert opinion that my photo shows a single egg at the tip of the female’s ovipositor.
Copyright © 2018 Walter Sanford. All rights reserved.
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Tags:adult female, adult male, American Bluets, contact guarding, Enallagma exsulans, Family Coenagrionidae, in tandem, mating pair, Narrow-winged Damselflies, ovipositor, Prince William Forest Park, South Fork Quantico Creek, Stream Bluet damselfly
Posted in Aperture, Canon 580EX Speedlite, damselflies, digital photography, education, natural science, Panasonic DMC-FZ150, photowalking, wildlife photography | Leave a Comment »
July 26, 2018
I’m honored to announce several of my dragonfly photographs are featured on new signage at Melvin L. Newman Wetlands Center, Clayton County, Georgia. The info-graphic, entitled “Mosquito Hawks,” was created by Danielle Bunch, Senior Conservationist for Clayton County Water Authority.

Image used with permission from Danielle Bunch.
As a retired K-12 science educator, I know from first-hand experience that informal learning opportunities can be as valuable as formal education in school classrooms. I was glad to contribute several of my photographs of Common Green Darner dragonflies (Anax junius) to the new info-graphic for the wetland area. It’s flattering to share the stage with Giff Beaton, author of Dragonflies and Damselflies of Georgia and the Southeast.
Full-size versions of my photographs (featured on the signage) appear in several previous posts on my photoblog.
Related Resources
Copyright © 2018 Walter Sanford. All rights reserved.
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Tags:adult female, adult male, Anax junius, Common Green Darner dragonfly, Family Aeshnidae (Darners), in tandem, in wheel, mating pair, oviposition
Posted in Aperture, Canon 580EX Speedlite, digital photography, dragonflies, education, natural science, Panasonic DMC-FZ150, wildlife photography | 4 Comments »
November 8, 2017
Two mating pairs of Autumn Meadowhawk dragonflies (Sympetrum vicinum) were photographed at Hidden Pond, Meadowood Recreation Area (MRA), Fairfax County, Virginia USA. Both pairs are “in tandem.”
The first pair is perching on the small wooden dock at Hidden Pond: the male is on the upper-right; the female is on the lower-left.
The last pair is perching on an American sweetgum tree (Liquidambar styraciflua) growing alongside the dock. I love the way the fall coloration of the tree leaves complements the coloration of the dragonflies! The male is on the upper-left; the female is on the lower-right.
Tech Tips
The photographs in this gallery were taken using my Panasonic Lumix DMC-FZ150 superzoom digital camera set for ~24x zoom (focal length of 600mm, 35mm equivalent), and Canon 580EX Speedlite external flash (manual mode).
In order to reduce “camera shake,” the camera was set for shutter priority mode. Using the reciprocal rule, the shutter speed was set for 1/800s. The ISO was set for “100.” An inexpensive Sunpak 6700M aluminum monopod was used for added stability.
Copyright © 2017 Walter Sanford. All rights reserved.
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Tags:adult female, adult male, Autumn Meadowhawk dragonfly, Family Libellulidae (Skimmers), gear talk, Hidden Pond, in tandem, mating pair, Meadowood Recreation Area, Sympetrum vicinum
Posted in Aperture, Canon 580EX Speedlite, digital photography, dragonflies, How To, natural science, Panasonic DMC-FZ150, wildlife photography | 1 Comment »
November 4, 2017
This blog post features more photos taken using my Fujifilm X-T1 digital camera, Fujinon XF55-200mm zoom lens, and a Fujifilm MCEX-11 extension tube. The camera was set for manual focus in order to use focus peaking; back-button focusing was used to focus automatically.
In wheel

ISO 640 | 200mm (~300mm, 35mm equivalent) | f/11 | 1/500s | 0.33 ev
Two of many mating pairs of Autumn Meadowhawk dragonflies (Sympetrum vicinum) were photographed on 27 October 2017 at Hidden Pond, Meadowood Recreation Area, Fairfax County, Virginia USA. Both pairs are “in wheel“: the male is on top; the female is on the bottom.

ISO 800 | 200mm (~300mm, 35mm equivalent) | f/18 | 1/500s | 0 ev
In tandem
The last mating pair is “in tandem“: the male is on the upper-right; the female is on the lower-left.

ISO 800 | 200mm (~300mm, 35mm equivalent) | f/16 | 1/500s | 0 ev
After copulation, Autumn Meadowhawks engage in a form of guarding behavior known as “contact guarding,” in which the male and female fly “in tandem” to egg-laying sites. Contact guarding is used by some species of odonates to prevent aggressive males from hijacking the female.
Related Resource: Adding an 11mm extension tube, a blog post by Walter Sanford.
Copyright © 2017 Walter Sanford. All rights reserved.
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Tags:adult female, adult male, Autumn Meadowhawk dragonfly, contact guarding, Family Libellulidae (Skimmers), gear talk, Hidden Pond, in tandem, in wheel, Meadowood Recreation Area, Sympetrum vicinum
Posted in Aperture, digital photography, dragonflies, education, extension tubes, Fujifilm X-T1, How To, natural science, wildlife photography | Leave a Comment »
August 20, 2017
A Powdered Dancer damselfly (Argia moesta) was spotted during a photowalk along a mid-size rocky stream in Fairfax County, Virginia USA. This individual is a male, as indicated by his terminal appendages. There is a whitish-blue morph female Powdered Dancer, therefore the male’s whitish-blue coloration is insufficient to identify its gender.
A week later, a mating pair of Powdered Dancers was spotted along the Potomac River at Riverbend Park, Fairfax County, Virginia USA. This pair is “in tandem“: the male is on the upper-left; the female on the lower-right.
The male is “contact guarding” the female as the pair flies “in tandem” to egg-laying sites where the female uses her ovipositor to insert eggs into vegetation (endophytic oviposition).
It’s helpful to take photos of mating pairs of damselflies, especially “in tandem,” since males and females of the same species can look quite different.
Female Powdered Dancers are polymorphic, including a whitish-blue andromorph and a brown heteromorph. The brown morph, shown in this pair, is more common than whitish-blue.
Did you notice the male Stream Bluet damselfly (Enallagma exsulans) perching near the Powdered Dancers? Thanks to Karen Kearney and Michael Boatwright, members of the Virginia Odonata Facebook group, for confirming my tentative identification of the Stream Bluet.
Copyright © 2017 Walter Sanford. All rights reserved.
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Tags:andromorph, Argia moesta, Fairfax County, heteromorph, in tandem, male, oviposition, ovipositor, polymorphic, Potomac River, Powdered Dancer damslelfly, Riverbend Park, terminal appendages, Virginia
Posted in Aperture, Canon 580EX Speedlite, damselflies, digital photography, natural science, Panasonic DMC-FZ150, photowalking, wildlife photography | 2 Comments »
December 24, 2016
More Big Bluet damselflies (Enallagma durum) were spotted in July 2016 during two photowalks along the Potomac River at Dyke Marsh Wildlife Preserve (DMWP).
More males
These individuals are males, as indicated by their blue and black coloration and by their terminal appendages.
Mating pair
The mating pair of Big Bluet damselflies shown in the following photograph is “in wheel,” in which the male uses “claspers” (terminal appendages) at the end of his abdomen to hold the female by her neck/thorax while they are joined at their abdomens. The male, blue and black in color, is on top; the female, green and black in color, is on the bottom.
The copulatory, or wheel, position is unique to the Odonata, as is the distant separation of the male’s genital opening and copulatory organs. Source Credit: Paulson, Dennis (2011-12-19). Dragonflies and Damselflies of the East (Princeton Field Guides) (Kindle Locations 377-378). Princeton University Press. Kindle Edition.
The wheel position is sometimes referred to as “in heart” when damselflies mate. In this case, the heart shape is deformed slightly.
The same pair is “in tandem” a while later: the male is on the right; the female is on the left. The male is engaged in “contact guarding,” in which the male and female fly “in tandem” to egg-laying sites. Contact guarding is used by some species of odonates to prevent aggressive males from hijacking the female.
Look closely at the underside of the female’s abdomen, near the tip. Notice the ovipositor that she uses to insert eggs into vegetation (endophytic oviposition).
It’s helpful to take photos of mating pairs of damselflies, especially “in tandem,” since males and females of the same species can look quite different.
Sidebar: Scientific Classification of Damselflies
The following concise explanation of the scientific classification of damselflies is provided to help the reader understand where the genus Enallagma (American Bluets) fits into the bigger picture of the Order Odonata, Suborder Zygoptera (Damselflies).
There are four families of damselflies in the United States of America, although only three families occur in the mid-Atlantic USA: Broad-winged damselflies; Narrow-winged damselflies (a.k.a., Pond Damselflies); and Spreadwing damselflies.
Family Calopterygidae is comprised of two genera.
Family Coenagrionidae is comprised of 14 genera. Three genera are common in Northern Virginia: Argia (Dancers); Enallagma (American Bluets); and Ischnura (Forktails).
- Argia (e.g., Blue-fronted Dancer, Blue-tipped Dancer, Variable Dancer)
- Enallagma (e.g., Big Bluet, Familiar Bluet, Orange Bluet, Stream Bluet)
- Ischnura (e.g., Eastern Forktail, Fragile Forktail, Rambur’s Forktail)
Family Lestidae is comprised of two genera.
- Archilestes (e.g., Great Spreadwing)
- Lestes (e.g., Slender Spreadwing, Southern Spreadwing, Swamp Spreadwing)
There are relatively few genera of Broad-winged Damselflies and Spreadwing Damselflies. In contrast, there are many more genera and species of Narrow-winged Damselflies — more species, including many that look similar, makes this family the most challenging to learn!
Related Resources: Excellent digital scans created by Gayle and Jeanelle Strickland. Click on the button labeled “Download file” in order to view full-size version of the graphics.
Copyright © 2016 Walter Sanford. All rights reserved.
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Tags:Big Bluet damselfly, claspers, contact guarding, Dyke Marsh Wildlife Preserve, Enallagma durum, Family Coenagrionidae, in heart, in tandem, in wheel, oviposition, ovipositor, Potomac River, terminal appendages
Posted in Aperture, Canon 580EX Speedlite, damselflies, digital photography, natural science, Panasonic DMC-FZ150, photowalking | Leave a Comment »
September 12, 2016
A mating pair of Dusky Dancer damselflies (Argia translata) was spotted along Pope’s Head Creek at Chapel Road Park. Dusky Dancer is a new species on my life list of odonates.
This pair is “in tandem“: the male is on the upper-right; the female is on the lower-left.
The male is on the upper-left in the following photo; the female is on the lower-right. The male is engaged in “contact guarding,” in which the male and female fly “in tandem” to egg-laying sites. The female is ovipositing in a partially submerged leaf.
Related Resource: A. translata male (Dusky Dancer) [JPG] [digital scan].
Copyright © 2016 Walter Sanford. All rights reserved.
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Tags:adult female, adult male, Argia translata, Chapel Road Park, Dancers (Argia), Dusky Dancer damselfly, Family Coenagrionidae, in tandem, mating pair, oviposition, Popes Head Creek, wildlife photography
Posted in Aperture, Canon 580EX Speedlite, damselflies, digital photography, natural science, Panasonic DMC-FZ150, photowalking | 1 Comment »
September 10, 2016
Two mating pairs of Powdered Dancer damselflies (Argia moesta) were spotted during a photowalk along Pope’s Head Creek at Chapel Road Park, Fairfax County, Virginia USA.
The first mating pair is “in heart“: the male is on top; the female is on the bottom.
Female Powdered Dancers are polymorphic; the female in the preceding mating pair is the blue morph that looks somewhat similar to males of the same species.
The last mating pair of Powdered Dancers was spotted “in tandem“: the male is on the upper-left; the female is on the lower-right. The female in the following mating pair is the tan morph.
Copyright © 2016 Walter Sanford. All rights reserved.
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Tags:adult female, adult male, Argia moesta, Chapel Road Park, Dancers (Argia), Family Coenagrionidae, in heart, in tandem, polymorphic, Popes Head Creek, Powdered Dancer damslelfly, wildlife photography
Posted in Aperture, Canon 580EX Speedlite, damselflies, digital photography, natural science, Panasonic DMC-FZ150, photowalking | Leave a Comment »
July 30, 2016
The Backstory: A cohort of emergent/teneral Slender Spreadwing damselflies (Lestes rectangularis) was discovered during late-May and early-June 2016 at a vernal pool located in Huntley Meadows Park (HMP). I have seen/photographed many female Slender Spreadwings in the past, but only one male. For the next few weeks, I focused upon finding and photographing mostly males from the cohort.
Mating Pair
A mating pair of Slender Spreadwing damselflies was spotted in a meadow located near a vernal pool from which the pair probably emerged. This pair is “in heart.”
All dragonflies and damselflies have a 10-segmented abdomen, numbered from front to back: male damselfly secondary genitalia, called hamules, are located in segments two and three (S2 and S3); female genitalia in segment eight (S8). Damselflies form the mating wheel (also known as the mating heart) in order for their genitalia to connect during copulation.
Therefore, the male is on upper-left; the female is on the lower-right.
The next photo shows the mating pair “in tandem,” immediately after copulation. Editor’s Note: Male (soft focus); female (sharp focus).
The last photo shows the mating pair, separated after being “in tandem.” The pair decoupled soon after the heart was broken. Editor’s Note: Male (sharp focus); female (soft focus).
Editor’s Note: This is Part 1 in a five-part series of blog posts documenting a cohort of Slender Spreadwing damselflies that emerged from a single vernal pool at Huntley Meadows Park, presented in reverse-chronological order from mature, reproducing adults to emergent tenerals.
Copyright © 2016 Walter Sanford. All rights reserved.
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Tags:adult female, adult male, claspers, Family Lestidae (Spreadwings), hamules, Huntley Meadows Park, in heart, in tandem, Lestes rectangularis, mating pair, Slender Spreadwing damselfly, teneral, terminal appendages, vernal pool, wildlife photography
Posted in Aperture, Canon 580EX Speedlite, damselflies, digital photography, education, natural science, Panasonic DMC-FZ150 | 6 Comments »